Each long strand of DNA combines with histones and nonhistone proteins to make up the substance chromatin.

每个长链DNA与组蛋和非组蛋构成染色物。

These results indicated that a nonhistone protein scaffold really existed and distributed as granules and fibres over the whole chromosomes.

染色体骨架在是真实存在的，骨架蛋以颗粒和纤维状的形式分布于整个染色体。