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In both types of feather, the barbules that extend from the barb ramus grow from a single layer of cells, called the barbule plate, on the periphery of the barb ridge.

而不论是何种出来的脊表面叫做「板」的单层细胞长出来。

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大英百科全书

Each barb branches into barbules, which link to one another by hooks.

每根羽支分拨成小羽,小羽通过钩子相互连接。

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Space – Verge Science

And lots of times, there are little feather barbules in there that we can look at the microscopic characters

很多时候, 里面有小羽,我们可以看到微观特征。

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PBS地球-动物趣闻

The central part of the feather became a hardened structure called a rachis, with lots of little barbs and barbules, creating the vanes of the feather.

中央部分变成了一种称为羽轴结构,上面有许多小倒钩和小枝,形成了羽叶片。

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